PLANT SWIMMING SPERM “requires an aqueous environment and thus limits their dispersal. Spermatophytes evolved mechanisms to protect their gametes from desiccation and so spread to almost all terrestrial areas. Flowering plants (angiosperms), which develop seeds from ovules that are enclosed in an ovary, represent the most successful phylum, comprising an extraordinary diversity of species dominating today’s terrestrial life.”
http://www.sciencedirect.com/science/article/pii/S1674205214608996