ADAPTATION TO DRY LAND. “Angiosperm fossil records dating back to the Lower Cretaceous (130–140 million years ago) show, in addition to the generation of seeds, the second major functional innovation of the spermatophytes: the evolution of pollen grains which harbor, and thus protect, sperm cells (the male gametes) from drying out.”
http://www.sciencedirect.com/science/article/pii/S1674205214608996