WIDESPREAD ANIMAL CULTURE. “Studies attribute additional orca behaviors, such as migration routes and song repertoires, to culture. Other research suggests that a finch’s song, a chimpanzee’s nut cracking, and a guppy’s foraging route are all manifestations of culture. Between 2012 and 2014, over 100 research groups published work on animal culture covering 66 species.”
http://www.pnas.org/content/114/30/7734.full