MODIFIED LAND NOSE. “Lungfish played a critical role when organisms shifted from an aquatic mode of life to a terrestrial setting representing one of the most dynamic major adaptive shifts during the course of evolution. The anatomy of lungfish shows that the olfactory passages open posteriorly in the oral region and into the respiratory portion of the organism. These posterior communicating pathways, however, were in all likelihood not used for the purposes of respiration but rather to increase the power of olfaction.”
http://www.wjgnet.com/esps/DownLoadFile.aspx?Type=Digital&SubType=2&DOI=10.5319%2Fwjo.v6.i2.33&FilePath=Pub%5C10.5319%5Cv6%5Ci2%5CWJO-6-33.doc