EXAPTATION OF GENES. “Whether megaphylls, which arose independently in four vascular plant lineages (ferns, sphenopsids, progymnosperms and seed plants), recruited the same gene systems is open to investigation. However, this does seem a possibility given that a common developmental mechanism for leaf production appears to have been recruited independently at least twice in the evolution of land plants.”
http://aob.oxfordjournals.org/content/96/3/345.full.pdf