MODIFYING GENES. “In species with genotypic sex determination, the chromosome pair that determines sex can change rapidly over time. Transitions are particularly likely when the ancestral sex chromosome exhibits little genetic differentiation, since WW or YY combinations are then less likely to be lethal. New sex-determining genes (or copies of the original gene in a new location) can lead to transitions within and between different XY and ZW system.”
http://journals.plos.org/plosbiology/article?id=10.1371/journal.pbio.1001899