CYTOPLASMIC REPLICONS. “The escaped element that would replicate in the cytoplasm using the ancestral Polinton pPolB spawned two groups of mobile elements, namely cytoplasmic plasmids (so far found only in fungi) and the “Megavirales,” which share the unique trifunctional capping enzyme, RNAP, and D11-like helicase.”
http://onlinelibrary.wiley.com/doi/10.1111/nyas.12728/full