METABOLIC ROLES. “Rubisco activity is downregulated during aging, in accordance with its postulated role as a nitrogen store. The observed profiles of sugars and methionine-related metabolites also suggest they play coordinated metabolic roles during ageing of maritime pine needles. Notably, pinitol is a ‘compatible solute’ involved in tolerance to osmotic and oxidative stresses in plants, and sucrose has cryoprotectant and osmoprotectant functions during overwintering.” (Rubisco, a major photosynthetic molecule).
http://jxb.oxfordjournals.org/content/66/11/3113.full