DEFAULT NIPPLES. “Darwin perpetuated the myth that male nipples are true evolutionary vestiges that stem from lactation in male as well as female ancestral mammals. They are, however, a developmental holdover from the early, indifferent stage of embryonic development, when the milk line arises but before the embryo has developed along the default female track or divergent male pathway. Male nipples are default structures formed earlier in development than the differentiation between males and females.”
https://evolution-outreach.springeropen.com/articles/10.1186/s12052-014-0012-5