SOCIAL NOT PHYLOGENETIC. “Neurobiological evidence also indicates that there are considerable socioecological variables involved in the evolution of facial displays among primate species. species that live in large, complex social groups had more facial nerve neurons than species that live in small social groups, indicating more potential control over mimetic musculature.”
http://onlinelibrary.wiley.com/doi/10.1111/joa.12440/full